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PACHYCEPHALIDAE - Whistlers1,2,3,4 (5:49)
MELANORECTES Sharpe, 1877 M - Rectes nigrescens Schlegel, 1871; type by monotypy5
Melanorectes nigrescens Black Whistler
nigrescens (Schlegel, 1871) Montane NW New Guinea (mountains of Doberai Pen.)
wandamensis (E. Hartert, 1930) vMontane NW New Guinea (mountains of Wondiwoi Pen.)
meeki (Rothschild & E. Hartert, 1913) Montane WC New Guinea (Maoke Mts.)
buergersi (Stresemann, 1922) Montane EC New Guinea (west to Victor Emanuel Range)
harterti Reichenow, 1911 Montane NE New Guinea (mountains of Huon Pen.)
schistaceus (Reichenow, 1900) vMontane SE New Guinea
CORACORNIS Riley, 1918 M - Coracornis raveni Riley, 1918; type by original designation   
Coracornis raveni   Riley, 1918 Maroon-backed Whistler
iMountains of C and SE Sulawesi
Coracornis sanghirensis 6  (Oustalet, 1881) Sangir Whistler
vSangir Is.
PACHYCEPHALA Vigors, 1825 F - Muscicapa pectoralis Latham, 1801; type by original designation   7
Pachycephala olivacea8 Olive Whistler
macphersoniana H.L. White, 1920 vSE Queensland, NE New South Wales
olivacea Vigors & Horsfield, 1827 vC and E Victoria, SE New South Wales
bathychroa Schodde & Mason, 1999 SC Victoria (Otway Range and Strzelecki Ranges) [Schodde, 1999 #3524]
apatetes Schodde & Mason, 1999 Tasmania, King I., Flinders I. [Schodde, 1999 #3524]
hesperus Schodde & Mason, 1999 SE South Australia, SW Victoria [Schodde, 1999 #3524]
Pachycephala rufogularis   Gould, 1841 Red-lored Whistler
vSC and SE South Australia, NW Victoria, SW and C New South Wales
Pachycephala inornata 9  Gould, 1841 Gilbert's Whistler
vS Western Australia to C New South Wales and NC Victoria
Pachycephala nudigula Bare-throated Whistler
ilsa Rensch, 1928 Sumbawa (Lesser Sundas)
nudigula E. Hartert, 1897 Flores (Lesser Sundas)
Pachycephala schlegelii Regent Whistler
schlegelii Schlegel, 1871 Montane NW Guinea (mountains of Vogelkop and Wondiwoi Pen.)
cyclopum E. Hartert, 193010 Montane N New Guinea (Cyclops Mts., ? Foya Mts.)
obscurior E. Hartert, 1896 vMontane N, WC, EC, NE and SE New Guinea (mountains of central cordillera and Huon Pen.; Torricelli Mts.)
Pachycephala soror Sclater's Whistler
soror P.L. Sclater, 1874 Montane NW New Guinea (mountains of Doberai Pen.)
octogenarii Diamond, 198511 Montane NW New Guinea (mountains of Bomberai Pen.) [Diamond, 1985 #1321]
klossi Ogilvie-Grant, 1915 Montane WC, EC, N and NE New Guinea (Kobowre Mts., Maoke Mts. and Central Highlands; Adelbert Range; mountains of Huon Pen.)
bartoni Ogilvie-Grant, 1915 Montane SE New Guinea (west to Herzog Range)
remota Mayr & Van Deusen, 1956 vMontane Goodenough I. (D'Entrecasteaux Arch.)
Pachycephala caledonica12 New Caledonian Whistler
vanikorensis Oustalet, 187513 αvVanikoro (Santa Cruz Is.)
intacta Sharpe, 1900 vBanks Is. to Efaté (Vanuatu)
chlorura G.R. Gray, 186014 αvErromango (Vanuatu)
cucullata (G.R. Gray, 1860)15 αvAnatom (Vanuatu)
littayei E.L. Layard, 1878 Ouvéa, Lifou (Loyalty Is.)
caledonica (J.F. Gmelin, 1789)16 vNew Caledonia, Ile des Pins
Pachycephala implicata Hooded Whistler
richardsi Mayr, 1932 Mountains of Bougainville (N Solomons)
implicata E. Hartert, 1929 vMountains of Guadalcanal (SE Solomons)
Pachycephala meyeri 17  Salvadori, 1890 Vogelkop Whistler
iMontane NW New Guinea (mountains of Doberai Pen.)
Pachycephala lorentzi   Mayr, 1931 Lorentz's Whistler
iMontane WC and EC New Guinea (E Maoke Mts. to Victor Emanuel Range)
Pachycephala modesta Brown-backed Whistler
telefolminensis Gilliard & LeCroy, 1961 vMontane EC New Guinea (W Central Highlands)
hypoleuca Reichenow, 1915 vMontane EC and NE New Guinea (E Central Highlands and mountains of Huon Pen.)
modesta (De Vis, 1894) vMontane SE New Guinea (west to Herzog Range)
Pachycephala tenebrosa 18  (Hartlaub & Finsch, 1868) Morning Whistler
vBabeldaob to Peleliu (Palau)
Pachycephala philippinensis Yellow-bellied Whistler
fallax (McGregor, 1904) Calayan (Philippines)
illex (McGregor, 1907) Camiguin Norte (Philippines)
philippinensis (Walden, 1872) vLuzon, Catanduanes (Philippines)
siquijorensis Rand & Rabor, 1957 vSiquijor (Philippines)
apoensis (Mearns, 1905) vSamar, Leyte, Mindanao (Philippines)
basilanica (Mearns, 1909) vBasilan (Philippines)
boholensis Parkes, 196619 vBohol (Philippines)
Pachycephala sulfuriventer 20  (Walden, 1872) Sulphur-bellied Whistler/Sulphur-vented Whistler
iMountains of Sulawesi
Pachycephala hypoxantha Bornean Whistler
hypoxantha (Sharpe, 1887) vMountains of Borneo (except Poi Range)
sarawacensis Chasen, 1935 vMountains of W Borneo (Poi Range in extreme W Sarawak)
Pachycephala cinerea21,22 Mangrove Whistler
cinerea (Blyth, 1847)23 δvNE India, Bangladesh, mainland SE Asia, Greater Sundas, W Lesser Sundas (Lombok) [Blyth, 1847 #368]
plateni (W. Blasius, 1888) Palawan (Philippines)
Pachycephala albiventris24 Green-backed Whistler
albiventris (Ogilvie-Grant, 1894)25 vN Luzon (Philippines)
crissalis (J.T. Zimmer, 1918) vC and S Luzon (Philippines)
mindorensis (Bourns & Worcester, 1894) vMindoro (Philippines)
Pachycephala homeyeri White-vented Whistler
homeyeri (W. Blasius, 1890) Mindanao, Sulu Arch. (Philippines), Siamil I. (off NE Borneo)
major (Bourns & Worcester, 1894)26 vCebu (Philippines)
winchelli (Bourns & Worcester, 1894) Tablas, Sibuyan, Masbate and Ticao to Panay and Negros (Philippines)
Pachycephala fulvotincta27 Fulvous-tinted Whistler
teysmanni Büttikofer, 1893 Salayar I. (off SW Sulawesi)
everetti E. Hartert, 1896 Tanahjampea I., Kalaotoa I., Madu I. (Flores Sea)
javana E. Hartert, 1928 vE Java, Bali
fulvotincta Wallace, 186428 vSumbawa to Alor (Lesser Sundas)
fulviventris E. Hartert, 1896 vSumba (Lesser Sundas)
Pachycephala phaionota 29  (Bonaparte, 1850) Island Whistler
vN Moluccas to islands in Banda Sea and Aru Is., Western Papuan Is., lowland NW New Guinea (extreme NW Vogelkop), small islands in Cenderawasih Bay
Pachycephala hyperythra Rusty Whistler
hyperythra Salvadori, 187630 αvLower montane NW and WC New Guinea (Vogelkop to S watershed of Maoke Mts., east to R. Fly)
sepikiana Stresemann, 1921 vLower montane N, WC and EC New Guinea (N watershed of Maoke Mts. and Central Highlands; North Coast Ranges; ? Foya Mts.)
reichenowi Rothschild & E. Hartert, 1911 Lower montane NE New Guinea (mountains of Huon Pen.)
salvadorii Rothschild, 1897 Lower montane EC, SC and SE New Guinea (S watershed west to Mt. Bosavi)
Pachycephala simplex31 Grey Whistler
rufipennis G.R. Gray, 1858 vKai Is. (SE Moluccas)
waigeuensis Stresemann & Paludan, 1932 vGebe (N Moluccas), Waigeo (Western Papuan Is.)
gagiensis Mayr, 1940 vGag (Western Papuan Is.)
griseiceps G.R. Gray, 1858 Batanta, Salawati, Misool, lowland NW Guinea (Vogelkop and neck), Aru Is.
miosnomensis Salvadori, 1879 vMios Num I. (in Cenderawasih Bay)
jobiensis A.B. Meyer, 1874 vLowland N New Guinea (Cenderawasih Bay to Astrolabe Bay), Yapen I.
perneglecta E. Hartert, 1930 vLowland SW to SE New Guinea (east to E Gulf of Papua)
brunnescens Wolters, 198032 Lowland NE and SE New Guinea (west to Huon Pen. and E Gulf of Papua), D'Entrecasteaux Arch. [Wolters, 1980 #4264]
sudestensis (De Vis, 1892) vTagula (Louisiade Arch.)
peninsulae E. Hartert, 1899 NE Australia (E Cape York Pen., NE Queensland)
simplex Gould, 1843 NC Australia (N Arnhem Land, Bathurst I., Melville I., Groote Eylandt)
Pachycephala arctitorquis33 Wallacean Whistler
kebirensis A.B. Meyer, 188434 αvRoma to Babar (Lesser Sundas)
arctitorquis P.L. Sclater, 1883 vTanimbar Is. (Lesser Sundas)
tianduana E. Hartert, 1901 vTayandu Is. (SE Moluccas)
Pachycephala orpheus 35  Jardine, 1849 Fawn-breasted Whistler
iSemau and Timor to Wetar (Lesser Sundas)
Pachycephala griseonota36 Drab Whistler
lineolata Wallace, 1863 vSula Is.
cinerascens Salvadori, 1878 Morotai, Halmahera, Ternate, Tidore, Bacan (N Moluccas)
johni E. Hartert, 1903 Obi (N Moluccas)
examinata E. Hartert, 1898 vBuru (S Moluccas)
griseonota G.R. Gray, 1862 Seram (S Moluccas)
kuehni E. Hartert, 1898 Kai Is. (SE Moluccas)
Pachycephala monacha37 Black-headed Whistler
monacha G.R. Gray, 1858 Aru Is. (off SW New Guinea)
dorsalis Ogilvie-Grant, 191138 vLocally in lowland and lower montane N, WC, EC, NE, SW and SE New Guinea [Salvadori, 1881 #3461]
Pachycephala leucogastra39 White-bellied Whistler
leucogastra Salvadori & D'Albertis, 187640 αvLowland SE New Guinea (west to Gulf of Papua)
meeki E. Hartert, 1898 Rossel (Louisiade Arch.)
Pachycephala rufiventris41,42 Rufous Whistler
minor F.R. Zietz, 1914 vMelville I. and Bathurst I. (Northern Territory)
falcata Gould, 184343 vNW and NC Australia (Kimberley Division, Arnhem Land)
pallida E.P. Ramsay, 187844 vNE Australia (E Gulf of Carpentaria to Cape York Pen.)
rufiventris (Latham, 1801)45,46 αvW, C, S and E Australia
xanthetraea (J.R. Forster, 1844) vNew Caledonia
Pachycephala lanioides White-breasted Whistler
carnarvoni (Mathews, 1913)47,48 δCoastal WC Western Australia (Pilbara region)
lanioides Gould, 1840 Coastal N Western Australia (W Kimberley Division)
fretorum De Vis, 1889 Coastal NE Western Australia, N Northern Territory and NW Queensland
Pachycephala orioloides49 Oriole Whistler
bougainvillei Mayr, 193250 Buka, Bougainville, Shortland (N Solomons)
orioloides Pucheran, 1853 Choiseul, Santa Isabel, Malakobi, Florida (C Solomons)
melanonota E. Hartert, 1908 vVella Lavella, Mbava, Ranongga (New Georgia Is., C Solomons)
centralis Mayr, 1932 vKolambangara to Nggatokae (New Georgia Is., C Solomons)
melanoptera Mayr, 1932 vRendova, Tetepare (New Georgia Is., C Solomons)
pavuvu Mayr, 1932 Pavuvu (SE Solomons)
sanfordi Mayr, 1931 Malaita (SE Solomons)
cinnamomea (E.P. Ramsay, 1879) vGuadalcanal (SE Solomons)
christophori Tristram, 1879 San Cristobal, Santa Ana (SE Solomons)
feminina Mayr, 1931 vRennell (S Solomons)
Pachycephala flavifrons   (Peale, 1848) Samoan Whistler
iSamoa
Pachycephala jacquinoti 51  Bonaparte, 1850 Tongan Whistler
iVava'u Group (Tonga) [Bonaparte, 1850 #407]
Pachycephala vitiensis52 Fiji Whistler/White-throated Whistler
ornata Mayr, 1932 vDuff Is., Swallow Is., Ndeni (Santa Cruz Is.)
utupuae Mayr, 1932 Utupua (Santa Cruz Is.)
graeffii Hartlaub, 1866 Waya, WC Viti Levu (Fiji)
optata Hartlaub, 1866 vE Viti Levu, Ovalau (Fiji)
kandavensis E.P. Ramsay, 1876 vBeqa, Kadavu and offshore islands (Fiji)
aurantiiventris Seebohm, 1891 vVanua Levu (except SE), Yaqanga (Fiji)
ambigua Mayr, 1932 vRabi, Kioa, SE Vanua Levu (Natewa Pen.) (Fiji)
torquata E.L. Layard, 1875 vTaveuni (Fiji)
koroana Mayr, 1932 vKoro (Fiji)
vitiensis G.R. Gray, 186053 αvGau (Fiji)
bella Mayr, 1932 vVatu Vara (Fiji)
lauana Mayr, 1932 vS Lau Group (Fiji)
Pachycephala citreogaster54 Bismarck Whistler
collaris E.P. Ramsay, 1878 vLouisiade Arch., (except Rossel), Egum Atoll and Alcester I. (off SE New Guinea)
rosseliana E. Hartert, 1898 vRossel (Louisiade Arch.)
citreogaster E.P. Ramsay, 1876 New Hanover, New Ireland, New Britain (Bismarck Arch.)
sexuvaria Rothschild & E. Hartert, 1924 vSt. Matthias Group (Bismarck Arch.)
goodsoni Rothschild & E. Hartert, 1914 Admiralty Is. (Bismarck Arch.)
tabarensis Mayr, 1955 vTabar (Tabar Is., Bismarck Arch.)
ottomeyeri Stresemann, 1933 Lihir I. (Hibernian Is.)
Pachycephala mentalis55 Black-chinned Whistler
tidorensis van Bemmel, 1939 vTernate and Tidore (N Moluccas)
mentalis Wallace, 1863 vMorotai, Halmahera, Bacan (N Moluccas)
obiensis Salvadori, 1878 vBisa and Obi (N Moluccas)
Pachycephala macrorhyncha56 Banda Sea Whistler
calliope Bonaparte, 185057,58 Semau to Timor and Wetar (Lesser Sundas)
sharpei A.B. Meyer, 188459 αBabar (Lesser Sundas)
dammeriana E. Hartert, 1900 vDamar (Lesser Sundas)
par E. Hartert, 1904 Roma (Lesser Sundas)
compar E. Hartert, 1904 Leti and Moa (Lesser Sundas)
fuscoflava P.L. Sclater, 1883 vTanimbar Is. (Lesser Sundas)
macrorhyncha Strickland, 184960 vAmbon and Seram (S Moluccas)
buruensis E. Hartert, 1899 vBuru (S Moluccas)
clio Wallace, 1863 Sula Is.
pelengensis Neumann, 1941 vBanggai Is.
Pachycephala pectoralis61 Australian Golden Whistler
balim Rand, 1940 Montane WC New Guinea (C Maoke Mts.)
pectoralis (Latham, 1801)62,63 αvE Queensland, NE and EC New South Wales
xanthoprocta Gould, 1838 vNorfolk I.
contempta E. Hartert, 1898 vLord Howe I.
youngi Mathews, 1912 SE Australia
glaucura Gould, 1845 vTasmania, King I., Flinders I.
fuliginosa Vigors & Horsfield, 182764,65 vSW and SC Australia (east to NW Victoria)
Pachycephala melanura66 Mangrove Golden Whistler
dahli Reichenow, 1897 Coastal SE New Guinea (west to Redscar Bay) and islets to E Bismarck Arch.
spinicaudus (Pucheran, 1853) Coastal SC and SE New Guinea (east to Redscar Bay), Torres Straits Is.
violetae Mathews, 191267 Coastal N Australia (Arnhem Land to SE Gulf of Carpentaria, Bathurst I., Melville I.)
melanura Gould, 184368 vCoastal NW Australia (Pilbara region to Kimberley Division)
robusta Masters, 1876 vCoastal NE Australia (W Cape York Pen. to NE Queensland; ? E Queensland)
Pachycephala aurea   Reichenow, 1899 Golden-backed Whistler
vLocally in lowland and lower montane N, WC, EC, NE and SE New Guinea
PSEUDORECTES Sharpe, 1877 M - Rectes ferrugineus Bonaparte, 1850; type by subsequent designation (Mathews, 1930, Systema Avium Australasianarum, p. 641).69
Pseudorectes ferrugineus Rusty Shrike-thrush
leucorhynchus (G.R. Gray, 1862) vWaigeo (Western Papuan Is.)
fuscus (Greenway, 1966) vBatanta (Western Papuan Is.)
brevipennis (E. Hartert, 1896) vAru Is. (off SW New Guinea)
ferrugineus (Bonaparte, 1850) vSalawati, Misool, lowland NW and SW New Guinea (Vogelkop to S foothills of Maoke Mts.)
holerythrus (Salvadori, 1878)70 vLowland N New Guinea (Cenderawasih Bay to upper R. Sepik and C north coast), Yapen I.
clarus (A.B. Meyer, 1894) vLowland N, NE, SE and SC New Guinea (west to lower R. Sepik and Astrolabe Bay in north)
Pseudorectes incertus 71  (van Oort, 1909) White-bellied Shrike-thrush
vLowland SW and SC New Guinea (upper R. Noord to upper R. Fly)
COLLURICINCLA Vigors & Horsfield, 1827 F - Colluricincla cinerea Vigors & Horsfield, 1827; type by monotypy = Turdus harmonicus Latham, 1801  72
Colluricincla tenebrosa Sooty Shrike-thrush
atra (Rothschild, 1931) vMontane WC New Guinea (N watershed of Maoke Mts.)
tenebrosa (Rothschild, 1911)73 vMontane WC and EC New Guinea (S watershed of Maoke Mts. to Victor Emanuel Range; Schrader Range)
Colluricincla megarhyncha74,75,76 Little Shrike-thrush
1 affinis (G.R. Gray, 1862) vWaigeo (Western Papuan Is.)
2 batantae (Meise, 1929) Batanta (Western Papuan Is.)
2 aruensis (G.R. Gray, 1858) vAru Is. (off SW New Guinea)
2 megarhyncha (Quoy & Gaimard, 1830)77 vSalawati, Misool, lowland and lower montane NW, WC and SW New Guinea (east to SE head of Cenderawasih Bay, S watershed of Maoke Mts. and R. Digul region)
3 goodsoni (E. Hartert, 1930)78 Lowland SC New Guinea (Trans-Fly region)
4 melanorhyncha (A.B. Meyer, 1874) vBiak I. (in Cenderawasih Bay)
4 obscura (A.B. Meyer, 1874)79 vLowland to lower montane N New Guinea (E Cenderawasih Bay to C north coast and entire Mamberamo basin including R. Taritatu, Van Rees and Foya Mts.), Yapen I.
5 maeandrina (Stresemann, 1921) vLowland to lower montane WC, EC and N New Guinea (N watershed of Maoke Mts. to Victor Emanuel Range and upper R. Sepik)
6 tappenbecki Reichenow, 1899 Lowland to lower montane N and EC New Guinea (C north coast to Astrolabe Bay, Sepik-Ramu basin and N watershed of Central Highlands)
6 madaraszi (Rothschild & E. Hartert, 1903) Lowland to lower montane NE New Guinea (Huon Pen.)
7 despecta (Rothschild & E. Hartert, 1903)80 vLowland to lower montane EC, SC and SE New Guinea (west to middle R. Fly and S watershed of Central Highlands)
8 fortis (Gadow, 1883) vD'Entrecasteaux Arch.
8 trobriandi (E. Hartert, 1896) Trobriand Is.
8 discolor De Vis, 1890 Tagula (Louisiade Arch.)
9 parvula Gould, 184581 vNW and NC Australia (Kimberley Division, Arnhem Land, Bathurst I., Melville I.)
10 aelptes Schodde & Mason, 1976 Coastal NE Northern Territory (SW Gulf of Carpentaria) [Schodde, 1976 #3502]
10 normani (Mathews, 1914) C and S Torres Strait Is., NE Australia (Cape York Pen. south-west to SE Gulf of Carpentaria)
10 griseata (G.R. Gray, 1858)82 vNE Queensland (R. Endeavour to Halifax Bay)
10 synaptica Schodde & Mason, 199983 vCoastal EC Queensland (Bowling Green Bay to N Broad Sound) [Schodde, 1999 #3524]
10 gouldii (G.R. Gray, 1858) Coastal EC Queensland (S Broad Sound to Hervey Bay)
10 rufogaster Gould, 1845 Coastal and subcoastal SE Queensland to NE New South Wales
Colluricincla boweri 84  E.P. Ramsay, 1885 Bower's Shrike-thrush
iNE Queensland
Colluricincla harmonica85 Grey Shrike-thrush
brunnea Gould, 184186 vNW and NC Australia (Kimberley Division, Arnhem Land, Bathurst I., Melville I., Groote Eylandt)
superciliosa Masters, 187687 vCoastal NE, SE and SC New Guinea (west to Markham-Ramu valley in north), Torres Strait Is., NE Australia (Cape York Pen.)
harmonica (Latham, 1801)88,89 αvNE to inland E, SE and SC Australia (south-west to Flinders Ranges, Yorke Pen. and Kangaroo I.)
strigata Swainson, 183890 αvTasmania, King I., Flinders I.
rufiventris Gould, 184191 vWC, SW, SC and C Australia (east to Eyre Pen. and west margin of Lake Eyre basin)
Colluricincla woodwardi 92  E. Hartert, 1905 Sandstone Shrike-thrush
iNW and NC Australia (Kimberley Division and Arnhem Land to SW Gulf of Carpentaria)

1 The Colluricinclidae previously made up of the genera Collurincla and Pitohui is largely subsumed here; but the pitohuis were non-monophyletic and the type species plus P. dichrous are transferred to the Oriolidae and P. cristatus moves to the Falcunculidae where it becomes Ornorectes cristatus. The two 'pitohuis' belong to the monotypic genera Melanorectes and Pseudorectes.
2 The genus Hylocitrea has been transferred to the Bombycillidae following Spellman et al. (2008) [Spellman, 2008 #11761]; Pachycare to the Acanthizidae following Norman et al. (2009) [Norman, 2009 #11985]; and following Schodde & Christidis (2014) [Schodde, 2014 #15669] Aleadryas to the Oreoicidae and Rhagologus to its own family.
3 Sequence of genera derived from Jønsson et al. (2010) [Jønsson, 2010 #12015], in conjunction with Jønsson et al. (2008) [Jønsson, 2008 #10613], Dumbacher et al. (2008) [Dumbacher, 2008 #11709] and Zuccon & Ericson (2012) (Zuccon, 2012) [Zuccon, 2012 #13486].
4 The two pitohuis belong to the monotypic genera Melanore and Pseudorectes.
5 Formerly included in Pitohui (now in the Oriolidae), but established as sister to Pachycephala and Coracornis by Jønsson et al. (2008, 2010) [Jønsson, 2008 #10613], [Jønsson, 2010 #12015], Dumbacher et al. (2008) [Dumbacher, 2008 #11709] and Zuccon & Ericson (2012) [Zuccon, 2012 #13486]. Melanorectes, Coracornis and Pachycephala form a genus group.
6 For treatment as a separate species from Colluricincla megarhyncha see Rozendaal & Lambert (1999) [Rozendaal, 1999 #3413], and placed in Coracornis following Jønsson et al. (2010) [Jønsson, 2010 #12015].
7 Pending the availability of molecular data, now in press, we retain the treatment of broad P. pectoralis split into the groups discerned by Galbraith (1956) [Galbraith, 1956 #1644] as used by Dickinson (2003) [Dickinson, 2003 #9533] as a response to too broad a taxon. But see Boles (2007) [Boles, 2007 #12231] for retention of a broad P. pectoralis with 59 subspecies.
8 For subspecies, see Schodde & Mason (1999) [Schodde, 1999 #3524] and Higgins & Peter (2002) [Higgins, 2002 #13592].
9 Includes gilbertii see Schodde & Mason (1999) [Schodde, 1999 #3524]. The spelling gilberti is an ISS.
10 Identity of population in Foya Mts unclear, see Mayr & Gilliard (1954) [Mayr, 1954 #7002] and Diamond (1972) [Diamond, 1972 #1318].
11 Population in Fakfak Mts. may represent another, undescribed form, see Diamond (1985) [Diamond, 1985 #1321].
12 This is group G. from Galbraith (1956) [Galbraith, 1956 #1644]; all forms previously part of a broad species P. pectoralis except caledonica, which was treated as a monotypic species by Mayr, 1967 [Mayr, 1967 #2561]. See also Boles (2007) [Boles, 2007 #12231].
13 This name appears in Vol. 2 of series 6 of this journal contra Mayr (1967) [Mayr, 1967 #2561] and does date from 1875.
14 For correct date see Sherborn (1934) [Sherborn, 1934 #3608] and Dickinson et al. (2011) [Dickinson, 2011 #13287].
15 For correct date see Sherborn (1934) [Sherborn, 1934 #3608] and Dickinson et al. (2011) [Dickinson, 2011 #13287].
16 For treatment as a monotypic species see Mayr (1967) [Mayr, 1967 #2561].
17 Tentative report from the Foya Mts. by Diamond (1985) not since confirmed, see Beehler et al. (2012) [Beehler, 2012 #14412].
18 Treated in the genus Pitohui by Mayr (1967) [Mayr, 1967 #2561], then mistakenly placed in the genus Colluricincla in Dickinson (2003) [Dickinson, 2003 #9533]. Since shown to be embedded in Pachycephala, where sister to P. meyeri, P. lorentzi and P. modesta by Jønsson et al. (2008, 2009) [Jønsson, 2008 #10801], [Jønsson, 2010 #12015] and Norman et al. (2009) [Norman, 2009 #11907].
19 For recognition see Dickinson et al. (1991) [Dickinson, 1991 #1361].
20 Including meridionalis see White & Bruce (1986) [White, 1986 #4194].
21 Treated as P. grisola by Dickinson (2003) [Dickinson, 2003 #9533] but see Walters (2003) [Walters, 2003 #4083] for establishment of applicability and precedence of P. cinerea. The subspecies treated follow Parkes (1989) [Parkes, 1989 #2941].
22 Considered to form a superspecies with P. albiventris and P. homeyeri, see Boles (2007) [Boles, 2007 #12231].
23 Includes vandepolli, butaloides and secedens, see Parkes (1989) [Parkes, 1989 #2941].
24 For treatment of this and P. homeyeri as separate species from P. cinerea see Parkes (1989) [Parkes, 1989 #2941]; but see Mayr (1967) [Mayr, 1967 #2561].
25 This name dates from June 1894 and takes precedence over the name mindorensis which dates from December.
26 For recognition see Dickinson et al. (1991) [Dickinson, 1991 #1361].
27 This is group A. from Galbraith (1956) [Galbraith, 1956 #1644]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
28 Includes jubilarii; see White & Bruce (1986) [White, 1986 #4194].
29 Includes stresemanni; see Mees (1965) [Mees, 1965 #2608].
30 For correct date of publication see Poggi (1996) [Poggi, 1996 #10153].
31 For a discussion of a potential split of a brown simplex group (including brunnescens) and a citrinoid griseiceps group see Schodde & Mason (1999) [Schodde, 1999 #3524].
32 Includes P. dubia Ramsay, 1879, preoccupied by Motacilla dubius Shaw, 1811 a junior synonymy of P. pectoralis (Latham, 1801), see Wolters (1980) [Wolters, 1980 #4264]. Ramsay's name was itself proposed to replace his own earlier name Eopsaltria brunnea Ramsay, 1877, which was preoccupied by Pachycephala brunnea Wallace, 1865.
33 For treatment as a separate species from P. rufiventris see Filewood & Peckover (1978) [Filewood, 1978 #9578] and Bruce in White & Bruce (1986) [White, 1986 #4194].
34 For correct date see Quaisser & Eck (2006) [Quaisser, 2006 #7425].
35 Includes wetterensis, see White & Bruce (1986) [White, 1986 #4194].
36 For treatment as a separate species from P. rufiventris see White & Bruce (1986) [White, 1986 #4194]. See also Boles (2007) [Boles, 2007 #12231].
37 For treatment as a separate species from P. rufiventris see Filewood & Peckover (1978) [Filewood, 1978 #9578], Bruce in White & Bruce (1986) [White, 1986 #4194] and Schodde & Mason (1999) [Schodde, 1999 #3524].
38 The name lugubris is said to take precedence over dorsalis Ogilvie-Grant, 1911, see Mees (1994) [Mees, 1994 #2636], and Dekker & Quaisser (2006) [Dekker, 2006 #9463]. However, as Salvadori (1881) [Salvadori, 1881 #3461] first placed this MSS name in synonymy Art. 11.6.1 of the Code (I.C.Z.N., 1999) [I.C.Z.N., 1999 #2059] requires the name to be attributed to him. This will apply only if the name lugubris w as used as valid before Mees noticed this.
39 For treatment as a separate species from P. rufiventris see Filewood & Peckover (1978) [Filewood, 1978 #9578] and Bruce in White & Bruce (1986) [White, 1986 #4194].
40 Poggi (2010) [Poggi, 2010 #12656] reported that the volume began to be despatched in March 1876. The date 1875 may well apply if a preprint can be located which confirms this.
41 Forms a superspecies with P. arctitorquis, P. griseonota, P. monacha and P. leucogastra; see Schodde & Mason (1999) [Schodde, 1999 #3524].
42 Circumscription of species and component Australian and New Caledonian subspecies follows Schodde & Mason (1999) [Schodde, 1999 #3524].
43 Implicitly includes colletti see Schodde & Mason (1999) [Schodde, 1999 #3524].
44 Implicitly includes dulcior see Schodde & Mason (1999) [Schodde, 1999 #3524].
45 Implicitly includes maudeae see Schodde & Mason (1999) [Schodde, 1999 #3524].
46 For reasons to revert to 1801 and not use 1802 see Schodde et al. (2010) [Schodde, 2010 #12422].
47 Implicitly includes bulleri see Schodde & Mason (1999) [Schodde, 1999 #3524].
48 Correct original spelling. The spelling carnaroni in Mayr (1967) [Mayr, 1967 #2561] and Dickinson (2003) [Dickinson, 2003 #9533] is an ISS.
49 This is group C. from Galbraith (1956) [Galbraith, 1956 #1644]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
50 Population named whitneyi and treated as a subspecies of P. melanura by Dickinson (2003) [Dickinson, 2003 #9533] now considered an interspecific hybrid; see Mayr & Diamond (2001) [Mayr, 2001 #10562].
51 Bonaparte's name has date precedence over Pachycephala melanops (Pucheran, 1853) see duPont (1976) [duPont, 1976 #1449].
52 This is group D. from Galbraith (1956) [Galbraith, 1956 #1644] with the addition of vitiensis (the oldest name) for geographical reasons by Dickinson (2003) [Dickinson, 2003 #9533]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
53 For correct date see Sherborn (1934) [Sherborn, 1934 #3608] and Dickinson et al. (2011) [Dickinson, 2011 #13287].
54 This is the Bismarck Arch. element of group H. from Galbraith (1956) [Galbraith, 1956 #1644]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
55 This is group B. from Galbraith (1956) [Galbraith, 1956 #1644]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
56 This is the Banda Sea element of group H. from Galbraith (1956) [Galbraith, 1956 #1644]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
57 Includes arthuri see White & Bruce (1986) [White, 1986 #4194].
58 Name dated 1851? by Mayr (1967) [Mayr, 1967 #2561] but see Zimmer (1926) [Zimmer, 1926 #4296].
59 For correct date see Quaisser & Eck (2006) [Quaisser, 2006 #7425].
60 Includes alfurorum see White & Bruce (1986) [White, 1986 #4194].
61 This is group F. from Galbraith (1956) [Galbraith, 1956 #1644] with balim added for zoogeographic reasons by Dickinson (2003) [Dickinson, 2003 #9533]; all forms previously part of a broad species P. pectoralis. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
62 Includes queenslandica and ashbyi see Schodde & Mason (1999) [Schodde, 1999 #3524].
63 For reasons to revert to 1801 and not use 1802 see Schodde et al. (2010) [Schodde, 2010 #12422].
64 Includes occidentalis see Schodde & Mason (1999) [Schodde, 1999 #3524].
65 May merit treatment as a separate species, see Jønsson et al. (2008) [Jønsson, 2008 #11227].
66 This is group E. from Galbraith (1956) [Galbraith, 1956 #1644] minus balim; all forms previously part of a broad species P. pectoralis. See also Galbraith (1967) [Galbraith, 1967 #15459]. But see Mayr (1967) [Mayr, 1967 #2561] and Boles (2007) [Boles, 2007 #12231].
67 This taxon was recognised by Galbraith (1956) [Galbraith, 1956 #1644], Mayr (1967) [Mayr, 1967 #2561] and by Nyári & Joseph (2013) [Joseph, 2013 #14878]. But see Ford (1983) [Ford, 1983 #1580] and Schodde & Mason (1999) [Schodde, 1999 #3524].
68 Includes bynoei and hilli see Schodde & Mason (1999) [Schodde, 1999 #3524].
69 Members of this genus formerly included in Pitohui (now in the Oriolidae), but recently established as monophyletic sister of Colluricincla by Jønsson et al. (2008, 2010) [Jønsson, 2008 #10613], [Jønsson, 2010 #12015], Dumbacher et al. (2008) [Dumbacher, 2008 #11709], Norman et al. (2009) [Norman, 2009 #11907] and Zuccon & Ericson (2012) [Zuccon, 2012 #13486]; Pseudorectes and Colluricincla thus form a genus group.
70 This subspecies and clarus intergrade broadly in central N and SC New Guinea, see Rand & Gilliard (1967) [Rand, 1967 #3199].
71 For inclusion in Pseudorectes as sister to P. ferrugineus, see Dumbacher et al. (2008) [Dumbacher, 2008 #11709].
72 Sequence of species derived from the phylogeny of Jønsson et al. (2010) [Jønsson, 2010 #12015].
73 The name Pachycephala tenebrosa Rothschild, 1911, was preoccupied in Pachycephala by Rectes tenebrosus Hartlaub & Finsch, 1868. This occurred in 2003 and now the two are not congeneric Art. 59.4 of the Code (I.C.Z.N., 1999) [I.C.Z.N., 1999 2059] requires reinstatement.
74 Despite deep DNA divergence and possible cryptic speciation, differentiation in this species complex unresolved due to on-going lineage sorting, introgression and incomplete sampling, see Deiner et al. (2011) [Deiner, 2011 #13217] and Nyári & Joseph (2013) [Nyári, 2013 #14878].
75 Rearrangements of subspecies limited here to groups suggested by them that are consistent morphologically and coherent geographically based on Ford (1979) [Ford, 1979 #15460].
76 For inclusion of C. parvula see Schodde & Mason (1976) [Schodde, 1976 #3502].
77 Includes misoliensis and ferrugineus; see Mees (1965, 1982) [Mees, 1965 #2608], [Mees, 1982 #2626], Ford (1979) [Ford, 1979 #15460] and Deiner et al. (2011) [Deiner, 2011 #13217].
78 Includes wuroi see Mees (1982) [Mees, 1982 #2626].
79 Includes hybridus and idenburgi; see Mees (1965, 1982) [Mees, 1965 #2608], [Mees, 1982 #2626], Ford (1979) [Ford, 1979 #15460] and Deiner et al. (2011) [Deiner, 2011 #13217].
80 Includes neos, superfluus and palmeri; see Mees (1965, 1982) [Mees, 1965 #2608], [Mees, 1982 #2626], Ford (1979) [Ford, 1979 #15460] and Deiner et al. (2011) [Deiner, 2011 #13217].
81 Includes conigravi see Schodde & Mason (1999) [Schodde, 1999 #3524].
82 Includes parvissima see Schodde & Mason (1999) [Schodde, 1999 #3524].
83 May prove consubspecific with gouldii see Nyári & Joseph (2013) [Nyári, 2013 #14878]; but see Schodde & Mason (1999) [Schodde, 1999 #3524].
84 Position in sequence reflects finding by Jønsson et al. (2010) [Jønsson, 2010 #12015] that this species is sister to C. harmonica, and not to C. megarhyncha.
85 Subspecies classification follows Schodde & Mason (1999) [Schodde, 1999 #3524].
86 Includes roebucki, parryi and julietae see Schodde & Mason (1999) [Schodde, 1999 #3524].
87 Includes tachycrypta see Schodde & Mason (1999) [Schodde, 1999 #3524].
88 Includes pallescens, anda and halmaturina see Schodde & Mason (1999) [Schodde, 1999 #3524].
89 For reasons to revert to 1801 and not use 1802 see Schodde et al. (2010) [Schodde, 2010 #12422].
90 For correct date see Dickinson et al. (2011) [Dickinson, 2011 #13287].
91 Includes whitei and kolichisi Ford, 1987 [Ford, 1987 #1584]; see Schodde & Mason (1999) [Schodde, 1999 #3524].
92 For monotypic treatment see Schodde & Mason (1999) [Schodde, 1999 #3524]; thus includes assimilis.
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