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SULIDAE - Gannets and Boobies1 (3:10)
PAPASULA Olson & Warheit, 1988 F - Sula abbotti Ridgway, 1893; type by monotypy  2
Papasula abbotti   (Ridgway, 1893) Abbott's Booby
iChristmas I. (Indian Ocean) >> Indian and SW Pacific Oceans
MORUS Vieillot, 1816 M - Fou de Bassan Buffon; type by monotypy = Pelecanus bassanus Linnaeus, 1758  3
Morus bassanus   (Linnaeus, 1758) Northern Gannet
vE Canada, Iceland, British Isles, W France, Heligoland, Norway
Morus capensis 4  (M.H.C. Lichtenstein, 1823) Cape Gannet
vIslands off coasts of Namibia and South Africa
Morus serrator 5  (G.R. Gray, 1843) Australasian Gannet
iS Australian coast, Tasmania, North Island (New Zealand)
SULA Brisson, 1760 F - Sula Brisson; type by tautonymy = Pelecanus leucogaster Boddaert, 1783  
Sula sula Red-footed Booby
sula (Linnaeus, 1766) Caribbean islands; tropical SW Atlantic (Fernando de Noronha I., Ilha de Trindade, Ascension I.)
rubripes Gould, 1838 Tropical Indian and Pacific Oceans east to Hawaiian Is., Line Is. (Kiritimati), Iles Marquises and Pitcairn Is.
websteri Rothschild, 1898 Islands off W Mexico (Isla Clarión, Isla San Benedicto, Islas Marías, Isla Isabela); Isla de Coco, Isla la Plata, Galapagos Is.
Sula leucogaster Brown Booby
leucogaster (Boddaert, 1783) Caribbean islands; islands off French Guiana (Iles du Salut) and Brazil (Atol das Rocas, Fernando de Noronha I. and Arch. de São Pedro e São Paulo south to Ilha de Alcatrazes); Cape Verde Is.; Ile Alcatraz (off Guinea); São Tomé, Príncipe, Annobón, Ascension I., St. Helena I.
brewsteri Goss, 18886 Islands in Golfo de California, Islas Revillagigedo, Islas Marías, Isla Isabela, Ile Clipperton
etesiaca Thayer & Bangs, 1905 vIslands in Golfo de Fonseca (Honduras) and in Golfo de Panamá; Isla de Coco; Isla Gorgona
plotus (J.R. Forster, 1844) S Red Sea, Indian Ocean, SE Asian seas to New Guinea and N Australia, W and C Pacific Ocean
Sula nebouxii Blue-footed Booby
nebouxii A. Milne-Edwards, 1882 Islands in Golfo de California, off W Mexico (Islas Marías), in Golfo de Fonseca (Honduras), in Golfo de Panamá, off S Colombia (Isla Gorgona), in Golfo de Guayaquil (Ecuador) and off N Peru (Islas Lobos de Tierra, Islas Lobos de Afuera)
excisa Todd, 1948 vGalapagos Is.
Sula variegata   (von Tschudi, 1843) Peruvian Booby
vOffshore islands and coast from N Peru (Piura) to C Chile (Isla Mocha)
Sula dactylatra Masked Booby
dactylatra Lesson, 1831 vBahamas, Caribbean islands; islands off NE Brazil (Atol das Rocas, Fernando de Noronha I., Arch. dos Abrolhos); Ascension I.
melanops Hartlaub, 1859 Red Sea to NW Indian Ocean
personata Gould, 18467 vE Indian Ocean (Cocos Is., Christmas I., Lesser Sundas, NC coast of Western Australia), Coral Sea (NE Australia) to Philippines, Micronesia, Polynesia and Hawaiian Is.; islands west of Mexico (Rocas Alijos, Isla Clarión, Isla San Benedicto, Ile Clipperton)
tasmani van Tets, Meredith, Fullagar & P.M. Davidson, 19888 Norfolk I., Lord Howe I., Kermadec Is. [van Tets, 1988 #14008]
Sula granti 9  Rothschild, 1902 Nazca Booby
iIsla Clarión, Isla San Benedicto, Ile Clipperton, Isla de Malpelo, Isla la Plata, Galapagos Is. >> Baja California to Ecuador10

1 Classification derived from Patterson et al. (2011) [Patterson, 2011 #12636]. Dorst & Mougin (1979) [Dorst, 1979 #1408] placed all species in Sula.
2 Sister to Morus + Sula; see Patterson et al. (2011) [Patterson, 2011 #12636] and references therein.
3 The species in Morus form a superspecies; see Dorst & Mougin (1979) [Dorst, 1979 #1408].
4 Sister species is M. serrator; see Patterson et al. (2011) [Patterson, 2011 #12636].
5 Includes rex; see Dorst & Mougin (1979) [Dorst, 1979 #1408].
6 Includes nesiotes; see Dorst & Mougin (1979) [Dorst, 1979 #1408].
7 Includes bedouti; see Dorst & Mougin (1979) [Dorst, 1979 #1408]. Also includes californica; see Pitman & Jehl (1998) [Pitman, 1998 #3094].
8 Includes fullagari O'Brien & Davies, 1990 [O'Brien, 1990 #2793]; see Steeves (2009) [Steeves, 2009 #11897] and Gill et al. (2010) [Gill, 2010 #12348].
9 For treatment as a separate species from S. dactylatra see Pitman & Jehl (1998) [Pitman, 1998 #3094].
10 The population on the Islas de los Desventurados may be this species (Pitman & Jehl 1998) [Pitman, 1998 #3094].
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