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BUCEROTIDAE - Hornbills1 (16:55)
BUCORVUS Lesson, 1830 M - Buceros abyssinicus Boddaert, 1783; type by monotypy  
Bucorvus abyssinicus   (Boddaert, 1783) Northern Ground Hornbill
vSenegal to NW Kenya, Ethiopia
Bucorvus leadbeateri 2  (Vigors, 1825) Southern Ground Hornbill
iS Kenya to SE and W DR Congo and Angola, south to N and E Botswana, E and SE South Africa
TOCKUS Lesson, 1830 M - Buceros erythrorhynchus Temminck, 1823; type by subsequent designation (G.R. Gray, 1840, A List of the Genera of Birds, p. 50).  
Tockus monteiri 3  Hartlaub, 1865 Monteiro's Hornbill
iSW Angola, N Namibia
Tockus erythrorhynchus4 Red-billed Hornbill
kempi Tréca & Érard, 2000 Senegal, Gambia, Guinea and W Mali [Tréca, 2000 #3874]
erythrorhynchus (Temminck, 1823) vE Mali east to Ethiopia and Somalia, south to NE Tanzania
ruahae A.C. Kemp & Delport, 20025 W and C Tanzania [Kemp, 2002 #2196]
ngamiensis Roberts, 19326,7 vNE Namibia, N Botswana, SW Zambia, N Zimbabwe
rufirostris (Sundevall, 1850)8 αvE Zambia, S Malawi to Zimbabwe (except NW), E Botswana, NE South Africa
damarensis (Shelley, 1888)9 vN Namibia
Tockus leucomelas10 Southern Yellow-billed Hornbill
elegans Hartlaub, 1865 SW Angola, NW Namibia
leucomelas (M.H.C. Lichtenstein, 1842)11 N and C Namibia to W Mozambique, Zimbabwe and N and E South Africa
Tockus flavirostris 12  (Rüppell, 1835) Eastern Yellow-billed Hornbill
vEritrea and Somalia to NE Tanzania
Tockus deckeni Von der Decken's Hornbill
deckeni (Cabanis, 1868)13 αSE Sudan, C Ethiopia, S Somalia, N and NE Kenya to C Tanzania [Cabanis, 1868 #5966]
jacksoni (Ogilvie-Grant, 1891)14 SE South Sudan, NE Uganda, NW Kenya
LOPHOCEROS Ehrenberg, 1828 M - Type by subsequent designation T. nasutus (see Peters 5: 254) or? L. melanoleucus (per Tentamen). SMSG TO DECIDE AND REVISE
Lophoceros fasciatus15 African Pied Hornbill
semifasciatus (Hartlaub, 1855) vS Senegal to S Nigeria
fasciatus (Shaw, 1812) αvSE Nigeria to N DR Congo, Uganda and N Angola
Lophoceros alboterminatus16 Crowned Hornbill
geloensis Neumann, 190517 vSW Ethiopia, Uganda and W Kenya to N Angola and N Zambia
alboterminatus Büttikofer, 188918 vS PR Congo to W Angola
suahelicus Neumann, 190519 vS Somalia and E Kenya to E and S Zambia, Mozambique and E South Africa (to Eastern Cape)
Lophoceros bradfieldi 20  (Roberts, 1930) Bradfield's Hornbill
S Angola, NE Namibia, N Botswana, S Zambia, W Zimbabwe
Lophoceros pallidirostris Pale-billed Hornbill
pallidirostris (Hartlaub & Finsch, 1870) vAngola, S DR Congo, W Zambia
neumanni Reichenow, 1894 SE Tanzania, Malawi, N Mozambique
Lophoceros nasutus African Grey Hornbill
nasutus (Linnaeus, 1766)21 vSenegal to Eritrea, N Uganda and W and C Kenya; SW Saudi Arabia, W Yemen
epirhinus (Sundevall, 1850)22 αvS Uganda, SE Kenya to Angola, C Namibia and N South Africa
Lophoceros hemprichii 23  (Ehrenberg, 1833) Hemprich's Hornbill
Ethiopia, Eritrea, NW Somalia, NW Kenya
Lophoceros camurus Red-billed Forest Hornbill
pulchrirostris (Schlegel, 1862)24 vSierra Leone to SW Ghana, SW Nigeria
camurus (Cassin, 1857) S Cameroon to W and NE DR Congo, W Uganda
BERENICORNIS Bonaparte, 1850 M - Buceros comatus Raffles, 1822; type by subsequent designation (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 84).  25
Berenicornis comatus   (Raffles, 1822) White-crowned Hornbill
vSW continental SE Asia, Thai-Malay Pen., Sumatra and Borneo
HORIZOCERUS Oberholser, 1899 M - 26
Horizocerus hartlaubi Black Dwarf Hornbill
hartlaubi (Gould, 1861) Sierra Leone to S Cameroon
granti (E. Hartert, 1895) WC to NE DR Congo, W Uganda
Horizocerus albocristatus Long-tailed Hornbill
albocristatus (Cassin, 1848) vSierra Leone to Ivory Coast
macrourus (Bonaparte, 1850) vGhana and Togo
cassini (Finsch, 1903) Benin, W Nigeria to Gabon, DR Congo and W Uganda
CERATOGYMNA Bonaparte, 1854 F - Buceros elatus Temminck, 1831; type by monotypy  
Ceratogymna atrata   (Temminck, 1835) Black-casqued Hornbill
vSierra Leone to SW Ghana; S Nigeria to DR Congo, SW South Sudan and NW Angola
Ceratogymna elata   (Temminck, 1831) Yellow-casqued Hornbill
vSW Senegal to Togo; S Nigeria to W Cameroon
BYCANISTES Cabanis & F. Heine, Sr., 1860 M - Buceros bucinator Temminck, 1824; type by monotypy  
Bycanistes fistulator Piping Hornbill
fistulator (Cassin, 1850) αSenegal to W Nigeria
sharpii (Elliot, 1873) SE Nigeria to Gabon and NW Angola
duboisi W.L. Sclater, 1922 C and N DR Congo to Central African Republic and W Uganda
Bycanistes bucinator   (Temminck, 1824) Trumpeter Hornbill
iSE Kenya, C Tanzania and S DR Congo to N and E Zimbabwe, Mozambique and E and SE South Africa
Bycanistes albotibialis 27  (Cabanis & Reichenow, 1877) White-thighed Hornbill
vBenin and S Nigeria to W Uganda and NW Angola
Bycanistes cylindricus   (Temminck, 1831) Brown-cheeked Hornbill
vSierra Leone to Ghana
Bycanistes subcylindricus Grey-cheeked Hornbill
subcylindricus (P.L. Sclater, 1871) αvSenegal and Gambia to Ghana; SW Nigeria
subquadratus Cabanis & Schütt, 188028 vSE Nigeria to N DR Congo, Uganda, NW Tanzania and W Kenya; Angola
Bycanistes brevis 29  Friedmann, 1929 Silvery-cheeked Hornbill
vW and S South Sudan, Ethiopia and Kenya to Malawi, E Zimbabwe and Mozambique
RHINOPLAX Gloger, 1841 F - Buceros scutata Boddaert, 1783; type by subsequent designation (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 84). = Buceros vigil J.R. Forster, 1781  
Rhinoplax vigil   (J.R. Forster, 1781) Helmeted Hornbill
iS Myanmar (S Tenasserim), Thai-Malay Pen., Sumatra, Borneo
BUCEROS Linnaeus, 1758 M - Buceros rhinoceros Linnaeus, 1758; type by subsequent designation (Elliot, 1882, Monogr. Bucerot., pt. 10, p. xvi).  
Buceros bicornis 30  Linnaeus, 1758 Great Hornbill
vHimalayas (east from Uttarakhand), SW and NE India, E Bangladesh, S Yunnan, mainland SE Asia, Sumatra
Buceros rhinoceros Rhinoceros Hornbill
rhinoceros Linnaeus, 175831 C and S Thai-Malay Pen., Sumatra
silvestris Vieillot, 1816 vJava
borneoensis Schlegel & S. Müller, 1840 vBorneo
Buceros hydrocorax Rufous Hornbill
hydrocorax Linnaeus, 1766 Luzon, Marinduque (Philippines)
semigaleatus Tweeddale, 1878 vPanaon, Bohol, Leyte, Samar (Philippines)
mindanensis Tweeddale, 187732 vBasilan, Mindanao (Philippines)
ANORRHINUS Reichenbach, 1849 M - Buceros galeritus Temminck, 1831; type by subsequent monotypy (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 83).  33
Anorrhinus galeritus 34  (Temminck, 1831) Bushy-crested Hornbill
vS Myanmar (S Tenasserim), Thai-Malay Pen., Sumatra, Natuna Is. (Natuna Besar), Borneo
Anorrhinus austeni 35,36  Jerdon, 1872 Godwin Austen's Brown Hornbill
NE India, SW China, N and E continental SE Asia
Anorrhinus tickelli   (Blyth, 1855) Tickell's Brown Hornbill
SW continental SE Asia, N Thai-Malay Pen.
OCYCEROS Hume, 1873 M - Buceros birostris Scopoli, 1786; type by subsequent designation (Elliot, 1882, Monogr. Bucerot., pt. 10, p. xvii).  37
Ocyceros griseus   (Latham, 1790) Malabar Grey Hornbill
vSW India
Ocyceros gingalensis 38  (Shaw, 1811) Sri Lanka Grey Hornbill
vSri Lanka
Ocyceros birostris   (Scopoli, 1786) Indian Grey Hornbill
vNE Pakistan, S Nepal, N and C India, NW Bangladesh
ANTHRACOCEROS Reichenbach, 1849 M - Buceros monoceros Shaw, 1811; type by subsequent monotypy (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 83). = Buceros coronatus Boddaert, 1783  
Anthracoceros coronatus 39  (Boddaert, 1783) Malabar Pied Hornbill
vC and SW India, Sri Lanka
Anthracoceros albirostris Oriental Pied Hornbill
albirostris (Shaw, 1808)40 vHimalayan foothills (east from Uttarakhand), NE India, Bangladesh, SW China, mainland SE Asia to C Thai-Malay Pen. [Shaw, 1808 #3593]
convexus (Temminck, 1832) αvS Thai-Malay Pen., Greater Sundas to Bali
Anthracoceros marchei   Oustalet, 1885 Palawan Hornbill
iPalawan Group (Philippines)
Anthracoceros malayanus 41  (Raffles, 1822) Black Hornbill
vC and S Thai-Malay Pen., Sumatra and SE satellites, Borneo
Anthracoceros montani   (Oustalet, 1880) Sulu Hornbill
iJolo, Tawi-Tawi (Sulu Arch.)
ACEROS Hodgson, 1844 M - Aceros nipalensis Hodgson, 1844; type by monotypy = Buceros nipalensis Hodgson, 1829  42
Aceros nipalensis 43,44  (Hodgson, 1829) Rufous-necked Hornbill
vFoothills of E Himalayas (west to Sikkim - formerly Nepal), NE India, SE Xizang, S Yunnan, W and N continental SE Asia
RHABDOTORRHINUS A.B. Meyer & Wiglesworth, 1898 M - 45
Rhabdotorrhinus leucocephalus   (Vieillot, 1816) Writhed Hornbill
vMindanao, Dinagat, Camiguin Sur (Philippines)
Rhabdotorrhinus waldeni 46  (Sharpe, 1877) Walden's Hornbill
Panay, Negros; formerly Guimaras (Philippines)
Rhabdotorrhinus corrugatus47 Wrinkled Hornbill
rugosus (Begbie, 1834)48 vC and S Thai-Malay Pen., Sumatra and NW satellites (Batu Is.) [Begbie, 1834 #246]
corrugatus (Temminck, 1832) vBorneo
Rhabdotorrhinus exarhatus Sulawesi Hornbill
exarhatus (Temminck, 1823) vN Sulawesi
sanfordi Stresemann, 1932 Sulawesi (except N), Muna I., Buton I.
PENELOPIDES Reichenbach, 1849 M - Buceros panini Boddaert, 1783; type by subsequent designation (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 84).  49
Penelopides panini50 Western Tarictic
†? ticaensis Hachisuka, 1930 vTicao (Philippines)
panini (Boddaert, 1783) Masbate, Panay, Negros (Philippines)
Penelopides affinis Southern Tarictic
samarensis Steere, 1890 vSamar, Leyte, Bohol (Philippines)
affinis Tweeddale, 1877 vDinagat, Mindanao (Philippines)
basilanicus Steere, 1890 vBasilan (Philippines)
Penelopides manillae Luzon Tarictic
manillae (Boddaert, 1783)51 δLuzon, Marinduque (Philippines)
subniger McGregor, 1910 vPolillo (Philippines)
Penelopides mindorensis   Steere, 1890 Mindoro Tarictic
vMindoro (Philippines)
RHYTICEROS Reichenbach, 1849 M - Buceros plicatus Latham, 1790; type by subsequent designation (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 84). = Buceros plicatus J.R. Forster, 1781  52
Rhyticeros cassidix 53  (Temminck, 1823) Knobbed Hornbill
Sulawesi and N and SE satellites
Rhyticeros subruficollis 54  (Blyth, 1843) Plain-pouched Hornbill
vW and SW continental SE Asia, N and C Thai-Malay Pen.
Rhyticeros narcondami   Hume, 1873 Narcondam Hornbill
iNarcondam (Andamans)
Rhyticeros plicatus55 Papuan Hornbill
plicatus (J.R. Forster, 1781) vAmbon, Seram (S Moluccas)
ruficollis (Vieillot, 1816) vN Moluccas, Western Papuan Is. (Waigeo, Batanta, Salawati), lowland NW, SW and SC New Guinea (east to head of Cenderawasih Bay in north and W Gulf of Papua in south)
jungei Mayr, 1937 Yapen I., lowland N and SE New Guinea (west to R. Mamberamo drainage in north and Gulf of Papua in south), D'Entrecasteaux Arch. (Goodenough I., Fergusson I.)
dampieri Mayr, 1934 New Britain, New Ireland, New Hanover (Bismarck Arch.)
harterti Mayr, 1934 Buka, Bougainville, Shortland, Fauro (N Solomons)
mendanae E. Hartert, 1924 Choiseul, Santa Isabel, New Georgia Is., Malaita, Guadalcanal (C and SE Solomons)
Rhyticeros undulatus 56  (Shaw, 1811) Wreathed Hornbill
vHimalayas (west to West Bengal), NE India, mainland SE Asia (except NE), Greater Sundas to Bali
Rhyticeros everetti   Rothschild, 1897 Sumba Hornbill
iSumba (Lesser Sundas)

1 Sequence of genera follows Gonzalez et al. (2013) [Gonzalez, 2013 #14679]. The Bucorvidae are subsumed herein.
2 Kemp & Crowe (1985) [Kemp, 1985 #2192] and others used the name cafer, but see Browning (1992) [Browning, 1992 #544].
3 Includes marjoriae; see Boix-Hinzen & Kemp in Hockey et al. (2005) [Hockey, 2005 #12724].
4 Treatment as a single broad species based on conflicting results of molecular studies, see Kemp & Delport (2002) [Kemp, 2002 #2196], Viseshakul et al. (2011) [Viseshakul, 2011 #12802] and Gonzalez et al. (2013) [Gonzalez, 2013 #14679].
5 For recognition see Kemp in Hockey et al. (2005) [Hockey, 2005 #12724].
6 For recognition see Dowsett et el. (2008) [Dowsett, 2008 #12725]. But see also Kemp in Hockey et al. (2005) [Hockey, 2005 #12724].
7 Original spelling nagmiensis; emended based on internal information.
8 Includes degens Clancey, 1964 [Clancey, 1964 #814]; see Kemp in Hockey et al. (2005) [Hockey, 2005 #12724].
9 For treatment as a separate species from T. erythrorhychus see Boix-Hinzen in Hockey et al. (2005) [Hockey, 2005 #12724].
10 For treatment as a separate species from T. flavirostris see Kemp (1988) [Kemp, 1988 #4396].
11 Includes parvior Clancey, 1959 [Clancey, 1959 #803]; see White (1965) [White, 1965 #4191].
12 Includes somaliensis; see Kemp (1988) [Kemp, 1988 #4396].
13 First described in 1868 in the Journal für Ornithologie on p. 360.
14 For treatment as a subspecies of T. deckeni; see Kemp (2001) [Kemp, 2001 #2195].
15 Considered to form a superspecies with T. alboterminatus and T. bradfieldi by Kemp (2001) [Kemp, 2001 #2195].
16 Subspecies classification follows White (1965) [White, 1965 #4191], but see also Kemp (2001) [Kemp, 2001 #2195].
17 Includes stegmanni; see Vincent (1934) [Vincent, 1934 #14200].
18 Includes angolensis; see Vincent (1934) [Vincent, 1934 #14200].
19 Includes australis; see White (1965) [White, 1965 #4191].
20 Implicitly includes williaminae; see Kemp in Hockey et al. (2005) [Hockey, 2005 #12724].
21 Includes forskali; see White (1965) [White, 1965 #4191].
22 Includes dorsalis Sanft, 1954 [Sanft, 1954 #3468]; see Kemp (1988) [Kemp, 1988 #4396].
23 Includes exsul; see Mackworth-Praed & Grant (1957) [Mackworth-Praed, 1957 #2465].
24 For recognition see White (1965) [White, 1965 #4191].
25 Retention of monotypic genus follows Wells (1999) [Wells, 1999 #4118], as now supported by Hübner et al. (2001) [Hübner, 2001 #2049] and Viseshakul et al. (2011) [Viseshakul, 2011 #12802].
26 For recognition and species composition see Gonzalez et al. (2013a, b) [Gonzalez, 2013 #14679], [Gonzalez, 2013 #14876], whose data allow Tropicranus to be subsumed here.
27 For treatment as a separate species from B. cylindricus see Sibley & Monroe (1990) [Sibley, 1990 #3636] and Chappuis (2000) [Chappuis, 2000 #13693].
28 For correction to authorship see Steinheimer & Dean (2007) [Steinheimer, 2007 #9522].
29 Includes omissus; see Mackworth-Praed & Grant (1937) [Mackworth-Praed, 1937 #14201].
30 Includes cavatus; see Kemp (1988) [Kemp, 1988 #2193]. This name was applied to the disjunct Western Ghats population, but even if recognised this name cannot be used; see Ripley (1961) [Ripley, 1961 #3310].
31 Includes sumatranus; see Chasen (1935) [Chasen, 1935 #708].
32 Includes basilanica; see Dickinson et al. (1991) [Dickinson, 1991 #1361].
33 Includes Ptilolaemus based on Gonzalez et al. (2013) [Gonzalez, 2013 #14679]. But see Rasmussen & Anderton (2005) [Rasmussen, 2005 #4749] and Viseshakul et al. (2011) [Visheshakul, 2011 #12802].
34 For treatment as a monotypic species see Kemp (1988) [Kemp, 1988 #2193]; thus includes carinatus and minor.
35 For separation from P. tickelli see Kemp (1995) [Kemp, 1995 #2194].
36 Includes indochinensis; see Kemp (2001) [Kemp, 2001 #2195].
37 Previously placed in Tockus but see Kemp (1995) [Kemp, 1995 #2194].
38 For treatment as a separate species from O. griseus see Kemp & Crowe (1985) [Kemp, 1985 #2192].
39 Synonyms presumably include both malabaricus and leucogaster, but types are probably not extant. Many records attached to these names belonged to albirostris, not coronatus; see Frith & Frith (1983) [Frith, 1983 #1615].
40 For restriction of authorship to Shaw see Dickinson et al. (2006) [Dickinson, 2006 #9496].
41 Includes deminutus Sanft, 1960 [Sanft, 1960 #3469], recognised by Smythies (1999) [Smythies, 1999 #3667] but not by Frith & Frith (1983) [Frith, 1983 #1615] or Wells (1999) [Wells, 1999 #4118].
42 For correction of authorship see Dickinson & Walters (2006) [Dickinson, 2006 #9468].
43 Implicitly includes yunnanensis Guan, 1989 [Guan Guanxun, 1989 #1837]; see Kemp (2001) [Kemp, 2001 #2195], although perhaps not evaluated.
44 For correct citation and date see Pittie (2012) [Pittie, 2012 #13832].
45 Recognition and species sequence based on Gonzalez et al. (2013) [Gonzalez, 2013 #14679]. Species recognition follows Kemp (2001) [Kemp, 2001 #2195].
46 For treatment as a separate species from R. leucocephalus see Kemp (1988) [Kemp, 1988 #2193] and Kennedy et al. (2000) [Kennedy, 2000 #2203].
47 Placed in this genus temporarily pending publication of a proposal for a monotypic genus due to the findings of Viseshakul et al. (2011) [Viseshakul, 2011 #12802].
48 For recognition see Sanft (1960) [Sanft, 1960 #3469]. Includes megistus; see van Marle & Voous (1988) [van Marle, 1988 #3920].
49 Seems to nest within Rhyticeros in Viseshakul et al. (2011) [Viseshakul, 2011 #12802]; however, see Gonzalez et al. (2013) [Gonzales, 2013 #14679] for basis for recognition along with Rhabdotorrhinus and Rhyticeros.
50 Treatment as consisting of a single polytypic species follows Kennedy et al. (2000) [Kennedy, 2000 #2203], but see also Kemp (1988, 1995) [Kemp, 1988 #2193] [Kemp, 1995 #2194]. Taxon sampling not yet sufficient to resolve reranking (Viseshakul et al. 2011) [Viseshakul, 2011 #12802].
51 Correct original spelling. Spelling manilloe in Peters (1945) [Peters, 1945 #3013] based on the later reprint not the original.
52 Species sequence based on Viseshakul et al. (2011) [Viseshakul, 2011 #12802] with R. corrugatus included but with a distinct lineage.
53 Includes brevirostris van Bemmel, 1951 [van Bemmel, 1951 #3910]; see White & Bruce (1986) [White, 1986 #4194].
54 For treatment as a species separate from R. plicatus see Kemp (1988) [Kemp, 1988 #2193]. See also Rasmussen (2000) [Rasmussen, 2000 #3217].
55 For alternative treatment as monotypic see Kemp (2001) [Kemp, 2001 #2195].
56 Includes ticehursti and the later aequabilis Sanft, 1960 [Sanft, 1960 #3649]; see Kemp (2001) [Kemp, 2001 #2195].
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