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OTIDIDAE - Bustards1 (11:26)
HETEROTETRAX Sharpe, 1894 M - Otis vigorsii A. Smith, 1831; type by original designation  2
Heterotetrax vigorsii3,4 Vigors's Bustard/Karoo Korhaan
namaqua Roberts, 19325 iS Namibia to W South Africa
vigorsii (A. Smith, 1831)6 iSW South Africa
Heterotetrax rueppelii Rüppell's Bustard/Rüppell's Korhaan
rueppelii (Wahlberg, 1856)7 δiNW Namibia, S Angola
fitzsimonsi Roberts, 1937 iWC Namibia [Roberts, 1937 #3359]
Heterotetrax humilis   (Blyth, 1855) Little Brown Bustardα
vE Ethiopia, N Somalia
LISSOTIS Reichenbach, 1848 F - Otis melanogaster Rüppell, 1835; type by subsequent designation (G.R. Gray, 1855, Cat. of the Genera and Subgenera of Birds, p. 109).  
Lissotis melanogaster Black-bellied Bustard
melanogaster (Rüppell, 1835) iSenegal to Ethiopia, south to S Angola and Zambia
notophila Oberholser, 1905 vZimbabwe, S Mozambique, E South Africa
Lissotis hartlaubii   (von Heuglin, 1863) Hartlaub's Bustard
iSE Sudan, E South Sudan and E Ethiopia to Kenya and NE Tanzania
ARDEOTIS Le Maout, 1853 F - Otis arabs Linnaeus, 1758; type by monotypy  8
Ardeotis arabs9 Arabian Bustard
†? lynesi (Bannerman, 1930) iW Morocco
stieberi (Neumann, 1907)10 iSenegal to E Sudan
arabs (Linnaeus, 1758) iNE Sudan, Eritrea, Ethiopia, NW Somalia, Djibouti, SW Saudi Arabia, W Yemen
butleri (Bannerman, 1930) iSouth Sudan, NW Kenya
Ardeotis kori Kori Bustard
struthiunculus (Neumann, 1907) iN Somalia and Ethiopia to Kenya and N Tanzania
kori (Burchell, 1822) iS Angola to Zimbabwe, S Mozambique and South Africa
Ardeotis nigriceps   (Vigors, 1831) Great Indian Bustard
iW and C India
Ardeotis australis 11  (J.E. Gray, 1829) Australian Bustard
vLowland SC New Guinea (Trans-Fly region), inland and N Australia
Ardeotis ludwigii   (Rüppell, 1837) Ludwig's Bustard
SW Angola, W Namibia to S South Africa
Ardeotis denhami12 Denham's Bustard
denhami (Children & Vigors, 1826)13 iSenegal, Gambia and SW Mauritania to W Ethiopia and N Uganda
jacksoni (Bannerman, 1930) iLower Congo basin, SW Angola, Kenya to N Botswana and NW Zimbabwe
stanleyi (J.E. Gray, 1831) iE and S South Africa [Gray, 1831 #1795]
Ardeotis heuglinii   (Hartlaub, 1859) Heuglin's Bustard
Eritrea, E and S Ethiopia, Djibouti, N and SC Somalia, N Kenya
Ardeotis nuba 14  (Cretzschmar, 1826) Nubian Bustard
W Mauritania to E Sudan
TETRAX T. Forster, 1817 M - Otis tetrax Linnaeus, 1758; type by tautonymy  
Tetrax tetrax 15  (Linnaeus, 1758) Little Bustard
iNW Africa and SW Europe to NW China >> Mediterranean basin to Pakistan
HOUBAROPSIS Sharpe, 1893 F - Otis bengalensis J.F. Gmelin, 1789; type by original designation  
Houbaropsis bengalensis Bengal Florican
bengalensis (J.F. Gmelin, 1789) vS Nepal to NE India
blandini Delacour, 1928 iS Cambodia, S Vietnam
SYPHEOTIDES Lesson, 1839 M - Otis aurita Latham, 1790; type by subsequent designation (G.R. Gray, 1840, A List of the Genera of Birds, p. 64). = Otis indica J.F. Miller, 1782  
Sypheotides indicus   (J.F. Miller, 1782) Lesser Florican
vPakistan, S Nepal, India
LOPHOTIS Reichenbach, 1848 F - Otis ruficrista A. Smith, 1836; type by monotypy  
Lophotis savilei 16  Lynes, 1920 Savile's Bustard
iSW Mauritania and Senegal to C Sudan
Lophotis gindiana 17  (Oustalet, 1881) Buff-crested Bustard
vEthiopia and Somalia to NE Tanzania
Lophotis ruficrista 18,19  (A. Smith, 1836) Red-crested Bustard
iS Zambia, S Angola and NE Namibia to S Mozambique, N South Africa, Swaziland
EUPODOTIS Lesson, 1839 F - Otis rhaad Rüppell, 1837; type by subsequent designation (G.R. Gray, 1840, A List of the Genera of Birds, p. 64). = Otis senegalensis Vieillot, 1820  
Eupodotis senegalensis20 White-bellied Bustard
senegalensis (Vieillot, 1820) vSW Mauritania and Guinea to C Sudan and Eritrea
canicollis (Reichenow, 1881)21 vEthiopia and Somalia to Uganda, N and E Kenya and NE Tanzania
erlangeri (Reichenow, 1905) iSW Kenya, W Tanzania
mackenziei C.M.N. White, 1945 iE Gabon, S DR Congo to W Zambia, C and S Angola [White, 1945 #4169]
barrowii (J.E. Gray, 1829) iE South Africa
Eupodotis caerulescens   (Vieillot, 1820) Blue Bustard/Blue Korhaan
iE South Africa (Mpumalanga to Free State and Eastern Cape)
OTIS Linnaeus, 1758 F - Otis tarda Linnaeus, 1758; type by subsequent designation (G.R. Gray, 1840, A List of the Genera of Birds, p. 64).  
Otis tarda Great Bustard
tarda Linnaeus, 175822 iSW and C Europe and NW Africa to Kazakhstan and W Xinjiang
dybowskii Taczanowski, 1874 iAltai Mts. and Mongolia to Transbaikalia and NE China
AFROTIS G.R. Gray, 1855 F - Otis afra Linnaeus, 1758; type by original designation  
Afrotis afra 23,24  (Linnaeus, 1758) Black Bustard/Southern Black Korhaanδ
vW and S South Africa
Afrotis afraoides White-quilled Bustard/Northern Black Korhaan
etoschae (Grote, 1922) iNW Namibia, NE Botswana
damarensis Roberts, 192625 vNamibia, W and C Botswana
afraoides (A. Smith, 1831)26 iSE Botswana and N and NE South Africa to Lesotho
CHLAMYDOTIS Lesson, 1839 F - Otis houbara Desfontaines, 1787; type by monotypy = Psophia undulata von Jacquin, 1784  
Chlamydotis undulata Houbara Bustard
fuertaventurae (Rothschild & E. Hartert, 1894) iE Canary Is.
undulata (von Jacquin, 1784) vN Africa east to Nile valley
Chlamydotis macqueenii 27,28  (J.E. Gray, 1832) Macqueen's Bustardδ
iS Ural Mts. to C Asia, Mongolia and N China, and SW Asia to W Pakistan >> N Africa to NW India, C China
1 Sequence of genera based on Pitra et al. (2002) [Pitra, 2002 #3096].
2 For recognition see Pitra et al. (2002) [Pitra, 2002 #3096], although type species (vigorsii) not sampled.
3 Forms a superspecies with H. rueppelii; see Snow (1978) [Snow, 1978 #13852].
4 The name scolopacea has not been used recently; it could be a prior name for namaqua. The type, in Leiden, needs to be reassessed.
5 Includes harei Roberts, 1937 [Roberts, 1937 #3359]; see White (1965) [White, 1965 #4191]. Also apparently includes barlowi Roberts, 1937 [Roberts, 1937 #3359]; see Hockey et al. (2005) [Hockey, 2005 #12724], but see also White (1965) [White, 1965 #4191].
6 Includes karrooensis Vincent, 1949 [Vincent, 1949 #4028]; see White (1965) [White, 1965 #4191].
7 Correct original spelling. No internal evidence permitting emendation.
8 Includes Neotis and Choriotis; see Pitra et al. (2002) [Pitra, 2002 #3096].
9 Considered to form a superspecies with A. kori by Collar (1996) [Collar, 1996 #1008].
10 Implicitly includes geyri Niethammer, 1954 [Niethammer, 1954 #2769]; see Vaurie (1965) [Vaurie, 1965 #3967] and Collar (1996) [Collar, 1996 #1008].
11 Considered to form a superspecies with A. nigriceps; see Collar (1996) [Collar, 1996 #1008].
12 Peters (1934) [Peters, 1934 #3007] used the name cafra for this species; for rejection of that name for this species see Meise & Stresemann (1950) [Meise, 1950 #14138]. Subsequently it was suppressed; see Opinion 407 (I.C.Z.N., 1956) [I.C.Z.N., 1956 #9627].
13 Includes burchellii; see Mackworth-Praed & Grant (1957) [Mackworth-Praed, 1957 #2465].
14 Includes agaze Vaurie, 1961 [Vaurie, 1961 #3964]; see Collar (1996) [Collar, 1996 #1008].
15 Includes orientalis; see Vaurie (1965) [Vaurie, 1965 #3967] and Cramp et al. (1980) [Cramp, 1980 #1084].
16 May not merit species rank; see Pitra et al. (2002) [Pitra, 2002 #3096].
17 Includes hilgerti; see White (1965) [White, 1965 #4191].
18 Includes ochrofacies Clancey, 1977 [Clancey, 1977 #880]; see Hockey et al. (2005) [Hockey, 2005 #12724].
19 Forms a superspecies with L. savilei and L. gindiana; see Sibley & Monroe (1990) [Sibley, 1990 #3636].
20 The name cafra Lichtenstein, 1793, used by Peters (1934) [Peters, 1934 #3007] for another species, is actually the earliest name for E. senegalensis; see Meise & Stresemann (1950) [Meise, 1950 #14138]. It has since been suppressed for priority; see Opinion 407 (I.C.Z.N., 1956) [I.C.Z.N., 1956 #9627].
21 Includes somaliensis; see Mackworth-Praed & Grant (1957) [Mackworth-Praed, 1957 #2465]. Also includes parva Moltoni, 1935 [Moltoni, 1935 #2699]; see White (1965) [White, 1965 #4191].
22 Includes korejewi; see Cramp et al. (1980) [Cramp, 1980 #1084].
23 Forms a superspecies with A. afraoides; see Sibley & Monroe (1990) [Sibley, 1990 #3636].
24 Correct original spelling. Spelling atra in Peters (1934) [Peters, 1934 #3007] an ISS. See Brooke & Dowsett (1969) [Brooke, 1969 #14231].
25 Includes mababiensis and kalaharica; see Mackworth-Praed & Grant (1962) [Mackworth-Praed, 1962 #4712]. Others see different bounds to these subspecies and place damarensis in afraoides, using kalaharica, for N and C Botswana birds see, e.g. Collar et al. (1986) [Collar, 1986 #14294].
26 Includes chiversi Roberts, 1933 [Roberts, 1933 #3358], and boehmeri Hoesch & Niethammer, 1940 [Hoesch, 1940 #1981]; see White (1965) [White, 1965 #4191]. Also includes leucoptera Reichenbach, 1848; see Roberts (1940) [Roberts, 1940 #13296].
27 For treatment as separate from C. undulata, see Gaucher et al. (1996) [Gaucher, 1996 #1690], Knox et al. (2002) [Knox, 2002 #2243], and Combreau et al. (2011) [Combreau, 2011 #13342]; thus treated here as allospecies.
28 Correct original spelling. Spelling mcqueeni in Dickinson (2003) [Dickinson, 2003 #9533] an ISS.
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